1. Department of Microbiology and Immunology, Hokkaido University Graduate School of Medicine, Kita-15, Nishi-7, Kita-ku Sapporo 060-8638, Japan
2. Department of Immunogy, Osaka Medical Center for Cancer, Nakamichi 1-3-2, Higashinari-ku, Osaka 537-8511, Japan
3. ERATO-JST, Laboratory of Molecular and Developmental Biology, University of Tsukuba, Tennoudai 1-1-1, Tsukuba 305-8577, Japan