1. Wolfson Centre for Mathematical Biology, Mathematical Institute, University of Oxford , Oxford , UK
2. Kennedy Institute of Rheumatology, University of Oxford , Oxford , UK
3. Pharmaceutical Sciences–Clinical Pharmacology, Roche Innovation Center Basel , Basel , Switzerland
4. Centre for Cancer Immunology, Institute for Life Sciences, University of Southampton , Southampton , UK
5. Centre for Immuno-oncology, Nuffield Department of Medicine, University of Oxford , Oxford , UK