1. Department of Human Genetics, Leiden University Medical Center , Einthovenweg 20, 2333 ZC Leiden , The Netherlands
2. Division of Genome Biology, Department of Medical Biochemistry and Biophysics, Science for Life Laboratory, Karolinska Institutet , SE-17177 Stockholm , Sweden
3. Department of Cell & Chemical Biology, Leiden University Medical Center , Einthovenweg 20, 2333 ZC, Leiden , The Netherlands
4. Novo Nordisk Foundation Center for Protein Research, University of Copenhagen , Blegdamsvej 3B, DK-2200 Copenhagen, Denmark
5. Institute of Biology Leiden, Leiden University , Sylviusweg 72, 2333 BE, Leiden , The Netherlands