1. Department of Cell and Developmental Biology , John Innes Centre, Norwich Research Park, Norwich NR4 7UH, UK
2. National Key Laboratory of Plant Molecular Genetics, CAS Center for Excellence in Molecular Plant Sciences, Institute of Plant Physiology and Ecology, Chinese Academy of Sciences , Shanghai , China
3. CAS-JIC Center of Excellence for Plant and Microbial Sciences (CEPAMS), Institute of Plant Physiology and Ecology, Chinese Academy of Sciences , Shanghai , China