1. CNRS, INSERM, Aix Marseille Univ, Institut Paoli-Calmettes, CRCM, Equipe labellisée par la Ligue Nationale contre le Cancer , Marseille , F-13009 , France
2. Department of Inflammation and Immunity, Lerner Research Institute, Cleveland Clinic Foundation , Cleveland , OH , USA
3. Department of Genetics and Genome Sciences, Case Western Reserve University , Cleveland , OH , USA