1. Terry Fox Laboratory , BC Cancer, Vancouver, British Columbia V5Z 1L3, Canada
2. Department of Medical Genetics, University of British Columbia , Vancouver, British Columbia V6H 3N1, Canada
3. Department of Biosciences, University of Oslo , 0316 Oslo, Norway
4. Cell and Developmental Biology Program, University of British Columbia V6T 1Z3 , Vancouver, British Columbia, Canada
5. School of Biomedical Engineering, University of British Columbia , Vancouver, British Columbia V6T 1Z3, Canada