1. College of Landscape Architecture and Forestry, Qingdao Agricultural University, Qingdao 266109, China
2. Shandong Peanut Research Institute, Shandong Academy of Agricultural Sciences, Qingdao 266100, China
3. Key Laboratory for Tobacco Gene Resources, Tobacco Research Institute, Chinese Academy of Agricultural Sciences, Qingdao 266101, China
4. The National Field Station of Liangzi Lake Ecosystem, Department of Ecology, College of Life Sciences, Wuhan University, Wuhan 430072, China