1. Center for Marine Environmental Studies (CMES), Ehime University, Matsuyama, Ehime 790-8577, Japan
2. Department of Bioscience and Bioinformatics, Kyusyu Institute of Technology, Iizuka, Fukuoka 820-0067, Japan
3. Graduate School of Environmental and Symbiotic Sciences, Prefectural University of Kumamoto, Kumamoto, Kumamoto 862-8502, Japan