1. Department of Biology, Howard University, 415 College Street NW, Washington, DC 20059, USA
2. Department of Biology, University of Hawaii, Hilo, 200 West Kawili Street, Hilo, HI 96720, USA
3. Department of Evolution and Ecology, University of California Davis, 1 Shields Avenue, Davis, CA 95616, USA
4. Friday Harbor Laboratories, University of Washington, Friday Harbor, WA 98250, USA
5. Department of Ecology and Evolutionary Biology, Cornell University, 215 Tower Road, Ithaca, NY 14853, USA