1. Biodesign Center, Key Laboratory of Engineering Biology for Low-carbon Manufacturing, Tianjin Institute of Industrial Biotechnology, Chinese Academy of Sciences , Tianjin 300308, PR China
2. College of Biotechnology, Tianjin University of Science and Technology , Tianjin 300457, PR China
3. University of Chinese Academy of Sciences , Beijing 101408, PR China
4. Haihe Laboratory of Synthetic Biology , Tianjin 300308, PR China