1. Graduate Group in Biochemistry and Molecular Biophysics, Perelman School of Medicine, University of Pennsylvania , Philadelphia , PA 19104 , USA
2. Department of Medicine, Perelman School of Medicine, University of Pennsylvania , Philadelphia , PA 19104 , USA
3. Graduate Group in Cell and Molecular Biology, Perelman School of Medicine, University of Pennsylvania , Philadelphia , PA 19104 , USA
4. Department of Cancer Biology, Perelman School of Medicine, University of Pennsylvania , Philadelphia , PA 19104 , USA