1. Infectious Diseases Translational Research Programme, Yong Loo Lin School of Medicine, National University of Singapore , Singapore 117545 , Singapore
2. Department of Biochemistry, Yong Loo Lin School of Medicine, National University of Singapore , Singapore 117596 , Singapore
3. Infection and Immunity Program, Biomedicine Discovery Institute and Department of Microbiology, Monash University , Clayton , Victoria , Australia
4. Department of Microbiology and Molecular Genetics, Institute for Medical Research Israel-Canada, Faculty of Medicine, The Hebrew University of Jerusalem , Jerusalem 9112102, Israel