1. School of Grassland Science, Beijing Forestry University , Beijing 100083 , China
2. National Hulunber Grassland Ecosystem Observation and Research Station, Institute of Agricultural Resources and Regional Planning, Chinese Academy of Agricultural Sciences , Beijing 100081 , China
3. Ministry of Agriculture Key Laboratory of Crop Nutrition and Fertilization, Institute of Agricultural Resources and Regional Planning, Chinese Academy of Agricultural Sciences , Beijing 100081 , China
4. Department of Biology, University of Western Ontario , London, ON , Canada
5. College of Grassland Science and Technology, China Agricultural University , Beijing 100083 , China
6. College of Resource Environment and Tourism, Capital Normal University , Beijing 100048 , China
7. College of Agro-grassland Science, Nanjing Agricultural University , Nanjing 210095 , China
8. Department of Biology, School of Arts and Sciences, National University of Mongolia , Ulaanbaatar 14200 , Mongolia