1. Nanjing Agricultural University College of Life Sciences, , No. 1 Weigang, Nanjing 210095, China
2. Nanjing Agricultural University State Key Laboratory of Crop Genetics and Germplasm Enhancement and the College of Horticulture, , No. 1 Weigang, Nanjing 210095, China
3. University of Connecticut Department of Plant Science and Landscape Architecture, , Storrs, CT 06269, USA