1. Department of Neurobiology, University of Pittsburgh School of Medicine , Pittsburgh, PA 15213 , USA
2. Department of Anatomy and Neurobiology, Program in Neuroscience, University of Maryland, School of Medicine , Baltimore, MD 21201 , USA
3. Eunice Kennedy Shriver National Institute of Child Health and Human Development, National Institutes of Health , Bethesda, MD 20892 , USA