1. Department of Clinical Neurosciences, University of Cambridge , Cambridge CB2 0QQ , UK
2. European Molecular Biology Laboratory, Genome Biology Unit , 69117 Heidelberg , Germany
3. Division of Computational Genomics and Systems Genetics, German Cancer Research Center (DKFZ) , 69120 Heidelberg , Germany
4. European Molecular Biology Laboratory, European Bioinformatics Institute , Cambridge CB10 1SD , UK
5. University of Cambridge, CRUK Cambridge Institute , Cambridge CB2 0RE , UK
6. Wellcome Sanger Institute , Wellcome Genome Campus, Cambridge CB10 1SA , UK
7. Department of Haematology, University of Cambridge , Cambridge CB2 0AW , UK
8. Wellcome-Medical Research Council Cambridge Stem Cell Institute, University of Cambridge , Cambridge CB2 0AW , UK