1. Graduate School of Natural Science and Technology, Shimane University , 1060 Nishikawatsu, Matsue, Shimane 690-8504 , Japan
2. Bioresource and Life Sciences, The United Graduate School of Agricultural Sciences, Tottori University , 4-101 Koyama-Minami, Tottori, Tottori 680-8553 , Japan
3. Institute of Agricultural and Life Sciences, Academic Assembly, Shimane University , 1060 Nishikawatsu, Matsue, Shimane 690-8504 , Japan