1. Department of Biochemistry and Molecular Biology, Dalhousie University, Halifax, Nova Scotia B3H 4R2, Canada
2. Population Genetics, Heinrich-Heine University D�sseldorf, D-40225 D�sseldorf, Germany
3. Canadian Institute for Advanced Research, Program in Integrated Microbial Biodiversity, Toronto, ON, Canada
4. CEPLAS – Cluster of Excellence in Plant Sciences, Heinrich-Heine University D�sseldorf, D-40225 D�sseldorf, Germany