1. Department of Environmental & Biological Sciences, University of Eastern Finland , PO Box 111, 80101 Joensuu , Finland
2. Department of Zoology, Faculty of Science, Sohag University , PO Box 82524, Sohag , Egypt
3. Department of Biological and Environmental Science, University of Jyväskylä , PO Box 35, 40014 Jyväskylä , Finland