1. Graduate School of Life Science, Tohoku University , 41 Kawauchi, Aoba, Sendai, Miyagi 980-0862, Japan
2. Center for Northeast Asian Studies, Tohoku University , Miyagi, Japan, 41 Kawauchi, Aoba, Sendai, Miyagi 980-0862, Japan
3. Biology Program, Faculty of Science, University of the Ryukyus , 1 Senbaru, Nishihara, Okinawa 903-0129 , Japan
4. Hahajima Branch of Ogasawara Ranger Office, The Ministry of the Environment, Shizukazawa, Hahajima , Ogasawara, Tokyo 100-2211, Japan