1. Department of Entomology, Michigan State University , 202 CIPS, 578 Wilson Road, East Lansing, MI 48824 , USA
2. U.S. Department of Agriculture – Agricultural Research Service, Pollinating Insect – Biology, Management, Systematics Research Unit Present Affiliation: , 1410 N. 800 E., Logan, UT 84341 , USA
3. Department of Entomology, University of Manitoba , 12 Dafoe Road, Winnipeg, MB R3T 2N2 , Canada
4. Program in Ecology, Evolutionary Biology, and Behavior, Michigan State University , East Lansing, MI 48824 , USA