1. Department of Biomolecular Engineering, University of California Santa Cruz, CA, 95064 USA
2. Department of Molecular, Cellular, Developmental Biology, University of California Santa Cruz, CA, 95064 USA
3. Department of Molecular and Cell Biology, School of Natural Sciences, University of California Merced, CA, 95340 USA
4. Institute for the Biology of Stem Cells, University of California Santa Cruz, CA, 95064 USA