1. Department of Biology, University of Virginia, Charlottesville, VA 22908, USA
2. Department of Microbiology, Immunology, and Cancer Biology, University of Virginia Health System, Charlottesville, VA 22908, USA
3. Division of Infectious Diseases and International Health, University of Virginia Health System, Charlottesville, VA 22908, USA
4. Department of Biochemistry and Molecular Genetics, University of Virginia Health System, Charlottesville, VA 22908, USA
5. Center for Public Health Genomics, University of Virginia, Charlottesville, VA 22908, USA