1. Department of Life Science, Center for Tropical Ecology and Biodiversity, Tunghai University , Taichung 40704 , Taiwan
2. Department of Entomology, National Taiwan University , Taipei 10617 , Taiwan
3. Department of Nature Resources and Environmental Studies, National Dong Hwa University , Hualien 97401 , Taiwan
4. Department of Zoology, Graduate School of Science, Kyoto University , Kyoto 606-8502 , Japan
5. Department of Life Science, National Taiwan Normal University , Taipei 11677 , Taiwan