1. Department of Zoology and Entomology, University of Queensland, QLD 4072 Australia
2. Centre de Biologie et de Gestion des Populations, Campus International de Baillarguet, 34980 Montferrier/Lez, France
3. Department of Mathematical Sciences, University of Aberdeen, Aberdeen AB24 3UE, United Kingdom
4. Museum of Vertebrate Zoology, University of California, Berkeley, California 94720-3160