1. Department of Ecology and Evolutionary Biology, University of California, Irvine, California 92697-2525
2. Institute of Marine Biology, Vladivostok 690041, Russia
3. Department of General Biology, Ecology and Soils, Far Eastern State University, Vladivostok 690600, Russia
4. Departament de Genética, Universitat Autónoma de Barcelona, 08193 Bellaterra (Barcelona), Spain