1. Hangzhou Normal University College of Life and Environmental Sciences, , Hangzhou 311121, China
2. Zhejiang University College of Agriculture and Biotechnology, , Hangzhou 310058, China
3. Chinese Academy of Agricultural Sciences Tea Research Institute, , Hangzhou 310008, China
4. University of Helsinki Department of Agricultural Sciences, Viikki Plant Science Centre, , P.O. Box 27, FI-00014, Finland