1. Zoological Museum, Biodiversity Unit, University of Turku, FI, Finland
2. Department of Biology, Faculty of Science, Ferdowsi University of Mashhad, Mashhad, Iran
3. Research Department of Zoological Innovations (RDZI), Institute of Applied Zoology, Faculty of Science, Ferdowsi University of Mashhad, Mashhad, Iran
4. Institute for Biological Problems of the North RAS, Magadan, Russia
5. Department of Zoology & Entomology, University of the Free State, Bloemfontein, South Africa