1. SynthSys and School of Biological Sciences, University of Edinburgh, Edinburgh, UK
2. Roslin Institute, University of Edinburgh, Roslin, UK
3. EPCC, Bayes Centre, University of Edinburgh, Edinburgh, UK
4. Laboratory for the Foundations of Computer Science, School of Informatics, University of Edinburgh, Edinburgh, UK
5. Institute for Adaptive and Neural Computation, School of Informatics, University of Edinburgh, Edinburgh, UK