1. Biodiversity Institute and Department of Ecology & Evolutionary Biology, University of Kansas , 1345 Jayhawk Blvd. Lawrence, Kansas 66045, United States
2. Department of Health Promotion and Policy, School of Public Health and Health Sciences, University of Massachusetts Amherst , 715 N Pleasant St, Amherst, Massachusetts 01003, United States
3. Fundacion Octaedro , Quito, Ecuador