1. Department of Forest and Conservation Sciences, University of British Columbia, Vancouver, British Columbia, V6T 1Z4, Canada
2. Pacific Biological Station, Fisheries and Oceans Canada, Nanaimo, British Columbia, V9T 6N7, Canada
3. Günther Analytics, 402-5775 Hampton Place, Vancouver, British Columbia, V6T 2G6, Canada
4. School of Resource and Environmental Management, Fisheries and Oceans Canada, Simon Fraser University, Burnaby, British Columbia, V5A 1S6, Canada
5. Department of Zoology and Faculty of Land and Food Systems, University of British Columbia, Vancouver, British Columbia, V6T 1Z4, Canada