1. Field Science Center for Northern Biosphere, Hokkaido University, Hakodate, 0400051, Japan
2. Wakkanai Fisheries Research Institute, Hokkaido Research Organization, Wakkanai, 0970001, Japan
3. Institute for East China Sea Research, Nagasaki University, Nagasaki, 8512213, Japan
4. Graduate School of Fisheries Sciences, Hokkaido University, Hakodate, 0418611, Japan