1. Department of Fisheries Science, Virginia Institute of Marine Science, William & Mary, Gloucester Point, VA 23062, USA
2. Australian Research Council Centre of Excellence for Coral Reef Studies, James Cook University, Townsville, QLD 4811, Australia
3. PSL Research University, EPHE-UPVD-CNRS, USR 3278 CRIOBE, Université de Perpignan, 58 Avenue Paul Alduy, Perpignan Cedex 66860, France
4. College of Science and Engineering, James Cook University, Townsville, QLD 4811, Australia
5. Centre for Sustainable Tropical Fisheries and Aquaculture, College of Science and Engineering, James Cook University, Townsville, QLD 4811, Australia