1. University of Oxford Department of Biology, , 11A Mansfield Road, Oxford OX1 3SZ, UK
2. University of Bristol School of Biological Sciences, , 24 Tyndall Avenue, Bristol BS8 1TH, UK
3. Institute of Conservation Science & Learning, Bristol Zoological Society , Clifton, Bristol BS8 3HA, UK
4. Université d’Antananarivo Zoologie et Biodiversité Animale, , BP 906 Antananarivo 101, Madagascar
5. Calgary Zoo Wilder Institute, , 1300 Zoo Road NE, Calgary, AB T2E 7V6, Canada