1. Guangdong Academy of Forestry Guangdong Provincial Key Laboratory of Silviculture, Protection, and Utilization, , 233 Guang Shan Yi Road, Tian He District, Guangzhou 510520, China
2. Guangdong Academy of Sciences Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Institute of Zoology, , 105 Xingang West Road, Haizhu District, Guangzhou 510520, China
3. Zunyi Normal University College of Biology and Agriculture, , Ping'an Avenue, Xinpu New District, Zunyi 563006, China