1. Oregon State University Oregon Cooperative Fish and Wildlife Research Unit, Department of Fisheries, Wildlife, and Conservation Sciences, , 104 Nash Hall, Corvallis, OR, 97331 USA
2. Geological Survey U.S. , Oregon Cooperative Fish and Wildlife Research Unit, Department of Fisheries, Wildlife, and Conservation Sciences, 104 Nash Hall, Corvallis, OR, 97331 USA
3. Geological Survey U.S. , Forest and Rangeland Ecosystem Science Center, 3200 SW Jefferson Way, Corvallis, OR, 97331 USA