1. Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China
2. Ministry of Education, Key Laboratory for Biodiversity Science and Ecological Engineering, College of Life Sciences, Beijing Normal University, Beijing 100875, China
3. University of Chinese Academy of Sciences, Beijing 100049, China
4. Center for Excellence in Animal Evolution and Genetics, Chinese Academy of Sciences, Kunming 650223, China