1. University of California Santa Cruz Institute of Marine Sciences, , 1156 High St., Santa Cruz, CA 95064, USA
2. Southwest Fisheries Science Center, National Marine Fisheries Service, National Oceanic and Atmospheric Administration Fisheries Ecology Division, , 110 McCallister Way, Santa Cruz, CA 95060, USA
3. University of Amsterdam Department of Theoretical and Computational Ecology, , Science Park 904, 1098 XH Amsterdam, The Netherlands