1. Centre for Regenerative Medicine , School of Pharmacy and Bioengineering, , Keele , UK
2. Keele University , School of Pharmacy and Bioengineering, , Keele , UK
3. Oswestry Keele Orthopaedic Research Group (OsKOR), The Robert Jones and Agnes Hunt Orthopaedic Hospital Foundation Trust , Oswestry , UK
4. School of Biosciences, Cardiff University , Cardiff , UK
5. School of Medicine, Keele University , Keele , UK