1. Epigenetics Institute, Department of Cell and Developmental Biology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA 19104, USA
2. Center of Excellence in Environmental Toxicology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA 19104, USA
3. Department of Orthopaedic Surgery, University of Michigan, Ann Arbor, MI 48109, USA
4. Department of Environmental Health Sciences, School of Public Health, University of Michigan, Ann Arbor, MI 48109, USA
5. Department of Environmental Medicine, University of Rochester School of Medicine and Dentistry, Rochester, NY14642, USA