1. Department of Microbiology and Infection Control and Prevention, Toho University Graduate School of Medicine , 5-21-16 Omori-nishi, Ota-ku, Tokyo 143-8540 , Japan
2. Department of Microbiology, School of Life and Environmental Science, Azabu University , Kanagawa , Japan
3. Department of Microbiology and Infectious Diseases, Toho University School of Medicine , Tokyo , Japan
4. Department of Bacteriology I, National Institute of Infectious Diseases , Tokyo , Japan
5. Antimicrobial Resistance Research Center, National Institute of Infectious Diseases , Tokyo , Japan