1. Department of Biology, Loyola University , Chicago, IL, 60660 , USA
2. Department of Science and Education, Integrative Research Center, Field Museum of Natural History , Chicago, IL, 60605 , USA
3. Department of Biological Sciences, The George Washington University , Washington, DC, 20052 , USA
4. Center for Integrative Biodiversity Discovery, Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science , Berlin, 10115 , Germany
5. Departments of Entomology and Ecology & Evolutionary Biology, Cornell University , Ithaca, NY, 14850 , USA