1. Department of Ecology and Evolutionary Biology, University of Connecticut , 75 N. Eagleville Rd., Storrs, CT 06269 , USA
2. Department of Integrative Biology, Oklahoma State University , 501 Life Sciences West, Stillwater, OK 74078 , USA
3. Department of Biological Sciences, University of Arkansas , 601 Science and Engineering, Fayetteville, AR 72701 , USA