1. Department of Ecology and Evolutionary Biology, Center for Ocean Health, University of California, 100 Shaffer Road, Santa Cruz, CA 95060, USA
2. Department of Ecology, Evolution, and Behavior, University of Minnesota, 1987 Upper Buford Circle, St Paul, MN 55108, USA
3. Department of Wildland Resources, Utah State University, 5230 Old Main Hill, Logan, UT 84322, USA
4. Department of Biology, Carleton College, 1 North College Street, Northfield, MN 55057, USA
5. College of Biological Sciences, 123 Snyder Hall, 1474 Gortner Avenue, Saint Paul, MN 55108, USA
6. Gombe Stream Research Center, PO Box 185, Kigoma, Tanzania