1. University of Exeter, School of Biosciences, Penryn Campus, Penryn TR10 9FE, UK,
2. FoAM - Kernow, Workshop E, Jubilee Warehouse, Commercial Road, Penryn TR10 8FG, UK,
3. University of Cambridge, Department of Zoology, Downing Street, Cambridge CB2 3EJ, UK, and
4. DST-NRF Centre of Excellence at the FitzPatrick Institute, University of Cape Town, Rondebosch 7701, South Africa