1. Institute of Biological and Environmental Sciences, University of Aberdeen, Zoology Building, Tillydrone Avenue, AB24 2TZ Aberdeen, UK,
2. Húsavik Research Center, University of Iceland, Hafnarstett 3, 640 Húsavik, Iceland, and
3. Marine Alliance for Science and Technology for Scotland, University of Aberdeen, Zoology Building, Tillydrone Avenue, AB24 2TZ Aberdeen, UK