1. Centre for Ecology and Conservation, University of Exeter, Penryn, Cornwall, UK
2. NERC Biomolecular Analysis Facility, Department of Animal and Plant Sciences, University of Sheffield, Sheffield, UK
3. School of Biology, Faculty of Biological Sciences, University of Leeds, Leeds, UK
4. Environment and Sustainability Institute, University of Exeter, Penryn, Cornwall, UK
5. National Wildlife Management Centre, Animal and Plant Health Agency, Gloucestershire, UK