1. Department of Animal and Plant Sciences, Alfred Denny Building, University of Sheffield, Western Bank, Sheffield S10 2TN, UK,
2. Max Planck Institute for Ornithology, Eberhard-Gwinner-Strasse, 82319 Seewiesen, Germany,
3. Evolution & Diversité Biologique, Bâtiment 4R1, Université de Toulouse Paul Sabatier, 118 Route de Narbonne, 31062 Toulouse Cedex 09, France, and
4. Department of Computer Science, Regent Court, University of Sheffield, 211 Portobello, Sheffield S1 4DP, UK