1. Institute of Systematics and Evolution of Animals, Polish Academy of Sciences , ul. Sławkowska 17, 31-016, Kraków , Poland
2. Department of Animal Ecology/Ecology and Genetics, Evolutionary Biology Centre, Uppsala University , Norbyvägen 18 D, 752 36 Uppsala , Sweden
3. Institute of Nature Conservation, Polish Academy of Sciences , Mickiewicza 33, 31-120 Kraków , Poland
4. Institute of Environmental Sciences, Jagiellonian University , Gronostajowa 7, 30-387 Kraków , Poland
5. Evolution and Ecology Research Centre, School of Biological, Environmental and Earth Sciences, University of New South Wales , Kensington 2052 , Australia