1. South African National Biodiversity Institute, Kirstenbosch Research Centre, Newlands, Cape Town, South Africa
2. Statistics in Ecology, Environment and Conservation, Department of Statistical Sciences, University of Cape Town, Rondebosch, Cape Town, South Africa
3. African Climate and Development Initiative, Department of Statistical Sciences, University of Cape Town, Rondebosch, Cape Town, South Africa
4. Evolutionary Biology and Molecular Biology, Department of Zoology and Entomology, Rhodes University, Grahamstown, South Africa
5. NRF Centre of Excellence at the Percy FitzPatrick Institute, University of Cape Town, Rondebosch, Cape Town, South Africa
6. Museum of Vertebrate Zoology and Department of Integrative Biology, University of California, Harmon Way, Berkeley, CA, USA